Repositorio Institucional de la Universidad de Oviedo

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This work is the result of the interaction between the INDICAGEOPAR project (MAGRAMA-727/12-OAPN) and GEOCAVE project (MAGRAMA-580/12-OAPN), both funded by the Spanish Autonomous Organism of National Parks (OAPN)-Ministry of Agriculture, Food Quality and Natural Environment. We thank the shepherds of Picos de Europa for their collaboration and to the speleological groups listed as Supplementary Data. The digital elevation model is courtesy of the Spanish National Geographic Institute.