Repositorio Institucional de la Universidad de Oviedo

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This study was conducted with permission of the Direccion General de Recursos Naturales of the Principado de Asturias. We acknowledge support from the Guarderias of the Medio Natural of the Patrulla Oso and the Fundacion Oso Pardo, as well as the Reserva Natural Integral de Muniellos and Parque Natural de las Fuentes del Narcea, Degana e Ibias. We thank J. Belant, R. Shideler, and 2 anonymous referees for their suggestions, which helped improve the manuscript; and A. Zarzo-Arias for Geographic Information System assistance.