Repositorio Institucional de la Universidad de Oviedo

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Funding was provided by the National Research Foundation (incentive funding for rated researchers awarded to E.Z.C. and F.D.), by the University of Pretoria (incentive funding for rated researchers awarded to E.Z.C. and F.D. and a research fellowship awarded to F.D.) and by the Spanish Ministry of Economy and Competitiveness (research fellowship awarded to F.D.). We are grateful to SAN Parks staff for administrative and logistic support.