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The authors are grateful to J. L. Acuña for his invitation to the BIOCANT3 oceanographic expedition (DOSMARES) where P. carboxydivorans M-227 was isolated. This study was financially supported by the Universidad de Oviedo (UNOV-11-MA-02) and Gobierno del Principado de Asturias (SV-PA- 13-ECOEMP-62). The polarimeter and IR and NMR equipment used in this work were acquired via grants for scientific and technological infrastructures from the Ministerio de Ciencia e Innovación [Grant Nos. PCT-010000-2010-4(NMR) and INP-2011-0016-PCT-010000 ACT6 (polarimeterand IR)]. We are also grateful to J. L. Caso and J. A. Guijarro for continuous support. This is a contribution of the Asturias Marine Observatory