Repositorio Institucional de la Universidad de Oviedo

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We thank C. Guardado, J. Rodrı´guez, R. Pe´rez, and D. O. Pastene for technical support and R. Lendrum for linguistic advice. We are also grateful to K. With and three anonymous reviewers who provided helpful suggestions to improve the manuscript. Field work was conducted with the permission of the Wildlife Service of Asturias. Funding was provided by the Spanish Government and the European Social Fund (FPI BES2009-25093 Grant to DM and CGL2008-01275, CGL2011-28430, and CGL2015-68963-C2-2-R Grants to DG).