E.F.P. received financial supportfrom the overnment of Asturias and the FP7–MarieCurie- COFUND programme of the European Commission (Grants ‘Clarín’ ACA14-19 and ACB17-19). H.W.P.acknowledges funding from the People Programme (MarieCurie Actions) of the European Union’s Seventh FrameworkProgramme FP7/2007-2013/ under REA grant agreementno. 607785 for the NASSTEC project. The Royal BotanicGardens, Kew, receive grant-in-aid from Defra.