This publication is an output of the projects financed by the Spanish Ministry of Economy, Industry and Competitiveness (AGL2014-54995-P and AGL2016-77633-P), the Government of Principado de Asturias (GRUPIN14-055), FEDER funding through COMPETE (Project: UID/AMB/50017/2013), and by National Funds through the Portuguese Foundation for Science and Technology (FCT) within the Project PTDC/AGRFOR/2768/2014. ME was supported by a fellowship from the Severo Ochoa Program (BP11117; Government of Principado de Asturias, Spain). JP was supported by a fellowship from the FPU (AP2010-5857; Ministry of Education, Spain). FCT (Fundação para a Ciência e a Tecnologia, Portugal) supported the fellowships of GP (SFRH/BPD/101669/2014). The Spanish Ministry of Economy and Competitiveness supported MM and LV through the Ramón y Cajal program (RYC-2014-14981 and RYC-2015-17871, respectively). FCT/MEC (Portugal) co-funding by the FEDER, within the PT2020 Partnership Agreement and Compete 2020 provide financial support to Centre for Environmental and Marine Studies (CESAM –UID/AMB/50017).