This research was supported by funds from INIA and FEDER cofunding through the projects: RTA 2010-00120-C32-01, RTA 2013-00048-C03-02. MO was supported by the projects M200871201 (Academy of Sciences of the Czech Republic) and LO1415 (Ministry of Education, Youth and Sports of the Czech Republic). LV was supported by MEYS CR Re. CZ.1.07/2.3.00/20.0256 and the Spanish Government through the Juan de la Cierva Programme (JCI-2012-12444). MM was funded by the Juan de la Cierva Programme (JCI-2011-08958) (Spanish Ministry of Economy and Competitiveness).